Johnstonellapunensis (Boraginaceae), a new species endemic to the dry Puna of Chile.

In an earlier molecular phylogenetic study, a sample of what was originally identified as Cryptanthahispida (Boraginaceae) from Chile, grouped with species of the genus Johnstonella . This sample was subsequently shown not to be C.hispida , but an undescribed species, endemic to the dry Puna of Chile. This new species is described here as Johnstonellapunensis , along with a key to all South American species of the genus. Johnstonellapunensis resembles other members of that genus in having an ovate fruit shape, ovate nutlets and a long style that extends beyond the nutlets. It is unusual in the genus in having a non-tuberculate, dimpled to rugulose nutlet surface sculpturing. Its closest relative within the genus is likely the South American J.diplotricha .