Beyond Bergmann's rule: Global variability in human body composition is associated with annual average precipitation and annual temperature volatility.

Beyond associations with mean annual temperature predicted by Bergmann's rule, variability in human body composition is also associated with mean annual temperature and inter-annual temperature volatility, with these associations further differing between hot and cold settings. Collectively, our results suggest that associations of human body composition with climate are complex for both physique (fat-free mass) and energy stores (adiposity).